I wrote that, so I'd better answer. Usual suspects = John de Lacy, Saher de Quincy, Richard & Gilbert de Clare, Roger & Hugh le Bigod.
Their descendants intermarried early and dozens of gateways are descended from all 6 as a bunch. Eg any trail that connects a gateway to most of the FitzAlans, Nevilles, Cliffords or Wentworths will score all 6 by branches that are already badged, or almost.
That's not a complete list of Bolles's barons, it's just the ones reached by the review line. The others will need work on other lines.
For Cudworth there's a message from April on my profile, dated May 9th 2015.