Thank you for all your answers. However, in the case of William Cottrell, Relationship Calculator only had to look for 14 generations, one less than 15. And the links between that ancestral profile and my own profile have been there for several years already. So none of the answers explain why this happens. Another odd thing is that I am descended from a Rev. John Crandall through four different lines, and yet i don't see my DNA information at his or any other Crandall profiles until his descendant, my 5th Great-Grandmother. Yet other DNA profiles are indicated at those very same ancestral profiles. The cause might be that descent from too many lines from the same ancestor somehow cancel each other out. PS. I just recalculated my relationship to William Cottrell using the 30 generations button, and it worked. It says that he is my 14th Great-Grandfather. Somehow i did not think that 14th Gr.Grandfather would be beyond 15 generations. But i was wrong! Thank you!