Johnston, George Harvey. "The Douglases of Angus." The Heraldry of the Douglases: With Notes on All the Males of the Family, Descriptions of the Arms, Plates and Pedigrees. Edinburgh: W. & A.K. Johnston, Limited, 1907.41. Print.
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It may be possible to confirm family relationships with David by comparing test results with other carriers of his Y-chromosome or his mother's mitochondrial DNA.
Y-chromosome DNA test-takers in his direct paternal line on WikiTree: